Early Warner Bros. are weird. None of the characters were well established, and their style would change by director. Something like that seems like Avery would have done it. They were all trained by Ub Iwerks, who felt that Disney had cornered the market on children's cartoons and had a focus on adult humored animation. Iwerks' ComiColor studio went out of business, in part because of his humane treatment of his animators and in part because the market he was geared towards didn't exist, but when Iwerks sold the business his team stayed together onto the Warner Brothers label, with Iwerks himself directing three cartoons for them before leaving to become an engineer at Disney. Warner was able to curtail the most inappropriate jokes, but the sick sensibility that Ub had given them was never fully purged.